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Yong, Hoi Sen (1968) Karyotype of Four Malayan Rats (Muridae, Genus Rattus Fischer). Cytologia, 33 (2). pp. 174-180. ISSN 0011-4545, DOI https://doi.org/10.1508/cytologia.33.174.
Yong, Hoi Sen (1969) Karyotypes of Three Species of Rats from Hong Kong and Thailand (Muridae, genus Rattus Fischer). Cytologia, 34 (3). pp. 394-398. ISSN 0011-4545, DOI https://doi.org/10.1508/cytologia.34.394.