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Yong, Hoi Sen (1969) Karyotypes of Malayan rats (Rodentia-Muridae, genus Rattus Fischer). Chromosoma, 27 (3). pp. 245-267. ISSN 0009-5915, DOI https://doi.org/10.1007/BF00326164.
Yong, Hoi Sen and Dhaliwal, S.S. (1972) Supernumerary (B-) chromosomes in the Malayan house rat, Rattus rattus diardii (Rodentia, Muridae). Chromosoma, 36 (3). pp. 256-262. ISSN 0009-5915, DOI https://doi.org/10.1007/BF00283244.