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Hashim, Rosli and Ito, F. and Gotoh, A. and Billen, J. (2009) Evolution of specialized spermatheca morphology in ant queens: insight from comparative developmental biology between ants and polistine wasps. Arthropod Structure & Development, 38 (6). pp. 521-525. ISSN 1467-8039, DOI https://doi.org/10.1016/j.asd.2009.08.003.
Hashim, Rosli and Ito, F. and Gotoh, A. and Billen, J. (2008) Comparison of spermatheca morphology between reproductive and non-reproductive females in social wasps. Arthropod Structure & Development, 37 (3). pp. 199-209. ISSN 1467-8039, DOI https://doi.org/10.1016/j.asd.2007.11.001.